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Atkinson, E.M., A.W. Bateman, L.M. Dill, M. Krkošek, J.D. Reynolds and S.C. Godwin. 2018. Oust the louse: leaping behaviour removes lice from wild juvenile sockeye salmon Oncorhynchus nerka. Journal of Fish Biology. doi: abs/10.1111/jfb.13684. | PDF

Phillips, JA*, SJ Peacock*, AW Bateman, M Bartlett, MA Lewis and M Krkošek. An asymmetric producer-scrounger game: body size and social foraging behavior of coho salmon. Theoretical Ecology. doi: 10.1007/s12080-018-0375-2 *Joint first authors. | PDF | CODE

Towers J., M.J. Halle, K. Symonds, G. Sutton, A. Morton, P. Spong, J.P. Borrowman, and J.K.B. Ford. 2018. Infanticide in a mammal-eating killer whale population. Scientific Reports 8, Article # 4366. | PDF


Kibenge M.J.T., Y. Wang, A. Morton, R. Routledge, and F.S.B. Kibenge. 2017. Formal comment on: Piscine reovirus: Genomic and molecular phylogenetic analysis from farmed and wild salmonids collected on the Canada/US Pacific Coast. PLoS ONE 12(11): e0188690. https://doi. org/10.1371/journal.pone.0188690. | PDF

Morton, A., R. Routledge, S. Hrushowy, M. Kibenge, and F. Kibenge. 2017. The effect of exposure to farmed salmon on piscine orthoreovirus infection and fitness in wild Pacific salmon in British Columbia, Canada. PLoS ONE 12 (12): e0188793. https://doi.org/10.1371/journal.pone.0188793. | PDF

Peacock, S.J., M.A. Lewis, M. Krkošek and S. Lele. 2017. Study design and parameter estimability for spatial and temporal ecological models. Ecology and Evolution 7: 762–770. doi: 10.1002/ece3.2618. | PDF

Peacock, S.J., Portner, L., Harrington, P.D., Kelly, G., Forbes, H., and C. Guinchard. 2017. Sea lice on juvenile wild salmon in the Broughton Archipelago, British Columbia. A report from the Salmon Coast Field Station Society. | PDF

Phillips, J.A.*, S.J. Peacock*, A.W. Bateman, M. Bartlett, M.A. Lewis and M. Krkošek. 2017. An asymmetric producer-scrounger game: body size and social foraging behavior of coho salmon. Theoretical Ecology doi: 10.1007/s12080-018-0375-2 *Joint first authors. | PDF


Bateman, A.W., S.J. Peacock, B. Connors, Z. Polk, D. Berg, M. Krkošek, and A. Morton. 2016. Recent failure to control sea louse outbreaks on salmon in the Broughton Archipelago, British Columbia. Canadian Journal of Fisheries and Aquatic Sciences: 1-9. doi: 10.1139/cjfas-2016-0122. | PDF

Groner, M.L.*, L.A. Rogers*, A.W. Bateman, B.M. Connors, L.N. Frazer, S.C. Godwin, M. Krkosek, M.A. Lewis, S.J. Peacock, E.E. Rees, C.W. Revie, and U.E. Schlaegel. 2016. Lessons from sea louse and salmon
epidemiology. Phil. Trans. R. Soc. B 371: 20150203.
http://dx.doi.org/10.1098/rstb.2015.0203. *Joint first authors. | PDF

Kibenge, M.J.T., T. Iwamoto, Y. Wang, A. Morton, R. Routledge, and F.S.B. Kibenge. 2016. Discovery of variant infectious salmon anaemia virus (ISAV) of European genotype in British Columbia, Canada, Virol. J. 13 17. | PDF

Morton, A., and R. Routledge R. 2016. Risk and precaution: Salmon farming. Marine Policy 74: 205-212. https://doi.org/10.1016/j.marpol.2016.09.022. | PDF

Peacock, S.J. 2016. Sea lice on juvenile wild salmon in the Broughton Archipelago, British Columbia. A report from the Salmon Coast Field Station Society. | PDF

Peacock, S.J., A.W. Bateman, M. Krkošek, B. Connors, S. Rogers, L. Portner, Z. Polk, C. Webb, and A. Morton. 2016. Sea-louse parasites on juvenile wild salmon in the Broughton Archipelago, British Columbia, Canada. Ecology 97 (7) pp.1887. | PDF

Peacock, S.J., A.W. Bateman, M. Krkošek and M.A. Lewis. 2016. The dynamics of coupled populations subject to control. Theoretical Ecology. doi:10.​1007/​s12080-016-0295-y. | PDF


Godwin, S., L. Dill, J. Reynolds, and M. Krkošek.  2015. Sea lice, sockeye salmon, and foraging competition: lousy fish are lousy competitors. Canadian Journal of Fisheries and Aquatic Sciences. 72(7), 113-120. | PDF | NEWS

Peacock, S.J., A.W. Bateman, M. Krkošek, and M.A. Lewis. 2015. Parasitism and food web dynamics of juvenile Pacific salmon. Ecosphere 6: art264. doi: 10.1890/ES15-00337.1 | PDF

Rees, E.E., S. St-Hilaire, S.R.M. Jones, M. Krkošek, S. DeDominicis, M.G.G. Foreman, T. Patanasatienkul, and C.W. Revie. 2015. Spatial patterns of sea lice infection among wild and captive salmon in western Canada. Landscape Ecology 30(6): 989-1004. | JOURNAL


Krkošek, M., C. Revie, B. Finstad, and C. Todd. 2014. Comment on Jackson et al. “Impact of Lepeophtheirus salmonis infestations on migrating Atlantic salmon, Salmo salar L., smolts at eight locations in Ireland with an analysis of lice-induced marine mortality” Journal of Fish Diseases. 37(4), 415-417. | JOURNAL

Peacock, S.J., B. Connors, M. Krkošek, J. Irvine, and M.A. Lewis. Can reduced predation offset negative effects of sea louse parasites on chum salmon? Proceedings of the Royal Society B: Biological Sciences. 281(1776), 20132913. | PDF


Kibenge M.J.T., T. Iwamoto, Y. Wang , A. Morton, R. Routledge, and F.S.B. Kibenge. 2013. Whole-genome analysis of piscine reovirus (PRV) shows PRV represents a new genus in family Reoviridae and its genome segment S1 sequences group it into two separate sub-genotypes, Virol. J. 10, 230–250. | PDF

Peacock, S.J., M. Krkošek, S. Proboszcz, C. Orr, and M.A. Lewis. 2013. Cessation of a salmon decline with control of parasites. Ecological Applications. 23(3), 606-620.

Rogers, L., S.J. Peacock, P. McKenzie, S. DeDominicis, S. Jones, P. Chandler, M. Foreman, C. Revie, and M. Krkošek. 2013. Modeling parasite dynamics on farmed salmon for precautionary conservation management of wild salmon. PLoS ONE. 8: e60096.

Thitiwan P., J. Sanchez, E.E. Rees, M. Krkošek, S.R.M. Jones, and C.W. Revie. 2013. Sea lice infestations on juvenile chum and pink salmon in the Broughton Archipelago, Canada from 2003 to 2012. Diseases of Aquatic Organisms. 105, 149-161.


Connors B.M., D.C. Braun, R.M. Peterman, A.B. Cooper, J.D. Reynolds, L.M. Dill, G.T. Ruggerone, and M. Krkošek. 2012. Migration links ocean-scale competition and local climate with exposure to farmed salmon to shape wild salmon dynamics. Conservation Letters.  5(4), 304-312.

Frazer, L.N., A. Morton, and M. Krkošek. 2012. Critical thresholds in sea lice epidemics: evidence, sensitivity, and subcritical estimation. Proceedings of the Royal Society B-Biological Sciences. 279(1735),1950-1958.


Ashander, J., M. Krkošek, and M. Lewis. 2011. Aquaculture-induced changes to dynamics of a migratory host and specialist parasite: a case study of pink salmon and sea lice. Theoretical Ecology. DOI 10.1007/s12080-011-0122-4

Connors, B., C. Lagasse, and L. Dill. 2011. What’s love got to do with it? Ontogenetic changes in drivers of dispersal in a marine ectoparasite. Behavioral Ecology. 22(3), 588-593.

Krkošek, M., and R. Hilborn. 2011. Sea lice (Lepeophtheirus salmonis) infestations and the productivity of pink salmon (Oncorhynchus gorbuscha) in the Broughton Archipelago, British Columbia, Canada. Canadian Journal of Fisheries and Aquatic Sciences. 68(1), 17-29.

Krkošek, M., B. Connors, A. Morton, M. Lewis, L. Dill, and R. Hilborn. 2011. Effects of parasites from salmon farms on wild salmon populations. Proceedings of the National Academy of Sciences of the USA. 108, 14700-14704.

Krkošek, M., B. Connors, P. Mages, S. Peacock, H. Ford, J. Ford, A. Morton, J. Volpe, R. Hilborn, L. Dill, and M. Lewis. 2011. Fish farms, parasites, and predators: Implications for salmon population dynamics. Ecological Applications. 21, 897-914.

Krkošek, M., R. Hilborn, R. Peterman, and T. Quinn. 2011. Cycles, stochastcity, and density dependence in pink salmon population dynamics. Proceedings of the Royal Society of London Series B. 278, 2060-2068.

Morton, A., A. McConnell, R. Routledge, and M. Krkošek. 2011. Sea lice dispersion and salmon survival in relation to fallowing and chemical treatment on salmon farms. ICES Journal of Marine Science. 68, 144-156.

Williams, R., M. Krkošek, E. Ashe, T. A. Branch, S. Clark, et al. 2011. Competing conservation objectives for predators and prey: estimating killer whale prey requirements for chinook salmon. PLoS ONE. 6(11).


Connors, B., M. Krkošek, J. Ford, and L.M. Dill. 2010. Coho salmon productivity in relation to direct and trophic transmission of sea lice from salmon aquaculture. Journal of Applied Ecology. 47, 1372-1377.

Connors B.M., B. Hargreaves, S.R.M. Jones, and L.M. Dill. 2010. Predation intensifies parasite exposure in a salmonid food chain. Journal of Applied Ecology.  47(6), 1365-1371

Krkošek, M. 2010. Host density thresholds and disease control for fisheries and aquaculture. Aquaculture Environment Interactions. 1, 21-32. (Invited Article for 1st Issue)

Krkošek, M. 2010. Sea lice and salmon in Pacific Canada: Ecology and policy. Frontiers in Ecology and the Environment. 8, 201-209.

Krkošek, M. and M.A. Lewis. 2010. An R0 theory for source-sink dynamics with application to Dreissena competition. Theoretical Ecology. 3, 24-43.

Krkošek, M., A.W. Bateman, S. Proboscsz, and C. Orr. 2010. Dynamics of outbreak and control of salmon lice on two salmon farms in the Broughton Archipelago. Aquaculture Environment Interactions. 1, 137-146.

Lewis, M.A., M. Krkošek, and M.J. Wonham. 2010. Dynamics of emerging wildlife disease. In S. Sivaloganathan (Ed) Mathematical Biology, Fields Institute Communications, American Mathematical Society.

Losos, C., J. Reynolds, and L.M. Dill. 2010. Sex-selective predation by threespine sticklebacks on sea lice: A novel cleaning behaviour. Ethology. 116.10, 981-989.

McConnell, A., R. Routledge, and B. Connors. 2010. The effect of artificial light on marine invertebrate and fish abundance in an area of active salmon farming. Marine Ecology Progress Series. 419:147-156.


Dill, L.M., C. Losos, B.M. Connors, and P. Mages. 2009. Comment on Beamish et al. 2005 “A proposed life history strategy for the salmon louse, Lepeophtheirus salmonis in the subarctic Pacific”. Aquaculture. 286, 154-155.

Keller, R., M.A. Lewis, D.M. Lodge, J.F. Shogren, and M. Krkošek. 2009. Putting bioeconomic research into practice. In Bioeconomics of Invasive Species: Integrating ecology, economics, policy and management. Editors: R. Keller, D.M. Lodge, M. A. Lewis and J. F. Shogren. Oxford University Press. Oxford.

Krkošek, M., A. Morton, J. Volpe, and M.A. Lewis. 2009. Sea lice and salmon population dynamics: Effects of exposure time for migratory fish. Proceedings of the Royal Society of London Series B. 276, 2819-2828.


Connors, B.M., M. Krkošek, and L.M. Dill. 2008. Sea lice escape predation on their host. Biology Letters. 4, 455-457.

Connors, B.M., E. Juarez-Colunga, and L.M. Dill. 2008. Effects of varying salinities on Lepeophtheirus salmonis survival on juvenile pink and chum salmon. Journal of Fish Biology. 72, 1825-1830.

Ford, J.S. and R.A. Myers. 2008. A global assessment of salmon aquaculture impacts on wild salmonids. PLoS Biology. 6(2), e33.

Krkošek, M., J. Ford, A. Morton, S. Lele, and M.A. Lewis. 2008. Response to comment on ‘Declining wild salmon populations in relation to parasites from farm salmon.’ Science. 322, 1790-1791.

Krkošek, M., J.S. Ford, A. Morton, S. Lele, and M.A. Lewis. 2008. Sea lice and pink salmon declines: A response to Brooks and Jones. Reviews in Fisheries Science. 16, 413-420.

Morton, A., R. Routledge, and M. Krkošek. 2008. Sea lice infestation of juvenile salmon and herring associated with fish farms off the east central coast of British Columbia. North American Journal of Fisheries Management. 28, 523-532.


Krkošek, M., J.S. Ford, A. Morton, S. Lele, R.A. Myers, and M.A. Lewis. 2007. Declining wild salmon populations in relation to parasites from farm salmon. Science. 318, 1772-1775.

Krkošek, M., A. Gottesfeld, B. Proctor, D. Rolston, C. Carr-Harris, and M.A. Lewis. 2007. Effects of host migration, diversity, and aquaculture on disease threats to wild fish populations. Proceedings of the Royal Society of London Series B. 274, 3141-3149.


Krkošek, M., M.A. Lewis, A. Morton, N. Frazer, and J. Volpe. 2006. Epizootics of wild fish induced by farm fish. Proceedings of the National Academy of Sciences of the USA. 103, 15506-15510.

Krkošek, M., M.A. Lewis, J. Volpe, and A. Morton. 2006. Fish farms and sea lice infestations of wild juvenile salmon in the Broughton Archipelago – A rebuttal to Brooks. Reviews in Fisheries Science. 14, 1-11.

Morton A. and R. Williams. 2006. Response of the sea louse, Lepeophtheirus salmonis, infestation levels on juvenile wild pink, Oncorhynchus gorbuscha, and chum, O. keta, salmon to arrival of parasitized wild adult pink salmon. Canadian Field Naturalist. 120(2), 199-204.

Morton, A.B. and R. Routledge. 2006. Mortality rates for juvenile pink and chum salmon (Oncorhynchus gorbuscha and keta) infested with sea lice (Lepeophtheirus salmonis) in the Broughton Archipelago. Alaska Fisheries Research Bulletin. 11(2), 146-152.

Morton A. and R.D. Routledge. 2006. Fulton’s condition factor: Is it a valid measure of sea lice impact on juvenile salmon? North American Journal of Fisheries Management. 26(1), 56-62.


Blaylock, R.B., R.M. Overstreet, and A. Morton. 2005. The pathogenic copepod Phrixocephalus cincinnatus (Copepoda: Pennellidae) in the eye of arrowtooth flounder, Atherestes stomias, and rex sole, Glyptocephalus zachirus, from British Columbia. Bulletin of the European Association of Fish Pathologists. 25(3), 116-123.

Ford, J.K.B., G.A. Ellis, D.R. Matkin, et al. 2005. Killer whale attacks on minke whales:Prey capture and antipredator tactics. Marine Mammal Science. 21(4), 603-618.

Krkosek, M., M.A. Lewis, and J.P. Volpe. 2005. Transmission dynamics of parasitic sea lice from farm to wild salmon. Proceedings of the Royal Society of London Series B. 272, 689-696.

Krkosek, M., A. Morton, and J.P. Volpe. 2005. Non-lethal assessment of juvenile pacific salmon for parasitic sea lice infections and fish health. Transactions of the American Fisheries Society. 134, 711-716.

Morton, A. and R.D. Routledge. 2005. Mortality rates for juvenile pink, Oncorhynchus gorbuscha, and chum, O. keta, salmon infested with sea lice, Lepeophtheirus salmonis, in the Broughton. Alaska Fisheries Research Bulletin. 11(2), 146-152.

Morton, A., R.D. Routledge, and R. Williams. 2005. Temporal patterns of sea louse infestation on wild Pacific salmon in relation to the fallowing of Atlantic salmon farms. North American Journal of Fisheries Management. 25(3), 811-821.

2004 & Previous

Morton, A., R.D. Routledge, C. Peet, et al. 2004. Sea lice (Lepeophtheirus salmonis) infection rates on juvenile pink (Oncorhynchus gorbuscha) and chum (Oncorhynchus keta) salmon in the nearshore marine environment of British Columbia, Canada. Canadian Journal of Fisheries And Aquatic Sciences. 61(2), 147-157.

Morton, A. and H. Symonds. 2002. Displacement of Orcinus orca (L.) by high amplitude sound in British Columbia, Canada. Journal of Marine Science. 59, 71-80.

Morton, A. and J. Volpe. 2002. Description of escaped farmed Atlantic salmon Salmo salar captures and their characteristics in one Pacific salmon fishery area in British Columbia, Canada, in 2000. Alaska Fisheries Research Bulletin. 9(2): 102-110.

Morton, A. 2000. Occurrence, photo-identification and prey of pacific white-sided dolphins (Lagenorhyncus obliquidens) in the Broughton Archipelago, Canada 1984-1998. Marine Mammal Science. 16(1), 80-93.

Morton, A. 1990. A quantitative comparison of the behaviour of resident and transient forms of the killer whale off the central British Columbia coast. Report of the International Whaling Commission. (Special issue 12), 245-248.

Morton, A. 1986. Sound and behavioral correlation in captive Orcinus orca. In: Kirkevold, B.C. and Lockhard, J.S. (eds.) Behavioral Biology of Killer Whales. Alan R. Liss, Inc. New York, pp. 303-333.


Adams, M. 2009. Factors mediating coho salmon (Oncorhynchus kisutch) distribution and abundance in freshwater systems of the Broughton Archipelago, B.C. Honours Thesis, School of Environmental Studies, University of Victoria.

Bartlett, M.C. 2017. Minihumps: Characterizing a coastal British Columbia kokanee population recently derived due to anthropogenic environmental change. Masters Thesis, Department of Ecology and Evolutionary Biology, University of Toronto. | PDF

Connors, B. 2011. Causes and consequences of dispersal in a marine parasite. Doctoral thesis, Department of Biological Sciences, Simon Fraser University

Ford, J. 2006. Demonstration of an impact of salmon farming on survival of wild salmonids in the North Atlantic and Pacific, using population dynamics data. Masters Thesis. Department of Biological Sciences, Dalhousie University.

Godwin, S. 2018. Sea lice in the North Pacific: from sub-lethal effects on wild salmon to parasite management and policy. Doctoral Thesis, Simon Fraser University

Krkošek, M. 2008. Effects of Salmon Aquaculture on Sea Lice & Salmon Population Dynamics. Doctoral Thesis, Department of Biological Sciences, University of Alberta.

Losos, C. 2008. Behavioural interactions of sea lice, threespine stickleback, and juvenile pacific salmon. Masters Thesis, Department of Biological Sciences, Simon Fraser University.

Mages, P. 2008. The effects of sea lice on juvenile pink salmon predation susceptibility. Masters Thesis, Department of Biological Sciences, Simon Fraser University.

McConnell, A. 2008. Honours Thesis, Department of Biological Sciences, Simon Fraser University.

Peacock, S.J. 2015. People, predators and parasites: Unexpected dynamics of sea lice in Pacific Canada. Doctoral Thesis, University of Alberta, Canada.

Schine, J. 2013. Listening to a Sense of Place: Acoustic Ethnography with Billy Proctor in the Broughton Archipelago, British Columbia. Masters Thesis, School of Communication, Simon Fraser University. | PDF online

Silvergeiter, M.P. 2009. Multiscale analysis of nest site selction and fledgling success by marbled murrelets (Brachyramphus marmoratus) in British Columbia.  Masters Thesis, Department of Biological Sciences, Simon Fraser University.


Dill, LM, M Krkosek, B Connors, SJ Peacock, AW Bateman, R Routledge, MA Lewis and J Reynolds. 2015. Critique of the Document “Information Regarding Concerns about Farmed Salmon – Wild Salmon Interactions.” 13 p. | PDF